Webster's English Dictionary

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subdominant

sub.dom.i.nant \.s*b-'da:m-(*-)n*nt, 's*b-\ n : something dominant to an 
   inferior or partial degree : as  : the fourth musical degree of the major 
   or minor scale (as F in the scal e of C) : an ecologically important 
   life-form subordinate in influence to the dom inants of a community - 
   subdominant aj